One fish, two fish

I like it when the author of the puzzle, this time David Bunker, incorporates a punny title inside the crossword after giving a list of themed clues. This puzzle had 4 themes and also two that actually intersect with the title in the center of the puzzle. This puzzles main clue: Arboreal rodent; Grizzly, eg; Earth's largest mammal; Western Raptor. A fun and colorful puzzle.